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The Tree of Life is the huge sacred tree about which Princess Chiamaka's Juma Tribe have built their village in their Tribal Lands. The tree is so wide at its base that the tribe has carved several entrances to its interior, incorporating it into their village life. It's giant roots break through the surface again long distances from the tree itself, providing natural bridges and archways in the landscape.
The tree is very special, indeed. Its roots reached down all the way into the Twilight World.
Because of Princess Chiamaka's pact with the Undead Archon, the tree now appears to be dying. Benedict claims that if it dies, the separation between life and death will end with it. One more thing for Scarlett's to do list.